Volume 110. Part 2 2005

CONTENTS

Ruchin A.B., Borkin L.J., Lada G.A., Litvinchuk S.N., Rosanov Yu.M., Ryzhov M.K. Morphological variation, genome size and population systems of the green frog (Rana escu-lenta complex) of Mordovia ... 3

KozJiechkin V.V., Shishikin A.S., Tumanov I.L. Some peculiarities of hunting of wolverine (Gulo gulo L.) for large ungulate animals in the forest zone of the Krasnoyarsk region ... 11

Mandelshtam M.Yu., Nikitsky N.B., Bibin A.R. The bark-beetles (Coleoptera, Scolytidae) from the subfamilies hylesininae and scolytinae (except tribus xyleborini, cryphalini, corthylini) of the western Caucasus ... 20

Sviridov A. V. Collections of the Zoological Museum of the Moscow Lomonossov State University: catalogue of the name-bearing types of lepidopteran taxa described after the year 1952 ... 31

Sotchivko A. V., Kaabak L. V. Butterflies from the Families Satyridae, Nymphalidae, Riodi-nidae and Lycaenidae (Lepidoptera, Rhopalocera) of the river basins of Oksu — Murgab and Pshart (Tadjikistan) ... 44

Scientific communications

Arkhipov V.Yu. Breeding biology of Brambling (Fringilla montifringilla 1.) in taiga of the Middle Yenisei ... 54

Grigoryeva O. V. Some problems of morphogenesis and season growth of the shoots of Draba sibirica (pall.) thell ... 58

Floristic notes ... 65

Critique and bibliography

Mirkin B.M. A book on the theory of biodiversity conservation. Rewiew on the book: Ye.A. Schwartz. «Biodiversity conservation: communities and ecosistems» ... 90

MORPHOLOGICAL VARIATION, GENOME SIZE AND POPULATION SYSTEMS OF THE GREEN FROG (RANA ESCULENTA COMPLEX) OF MORDOVIA

A.B. Ruchin, L.J, Borkin, G.A. Lada, S.N. Litvinchuk, Yu.M. Rosanov, M.K. Ryzhov

The genome size in hybridogenic Rana esculenta is intermediate between that of parental species, differed in 7,5% (R. lessonae) and 7,2% (R. ridibunda), respectively. In Mordovia, six kinds of population systems were revealed. Two kinds of so-called «pure» (one-species) systems are most widely distributed. Three kinds of mixed population systems with participation of R. esculenta were identified: LE (3 localities), RE (1 locality), REL (6 localities). Relatively high occurrence of LR system may be explained by problems with exact species identification of green frogs in the field based on external morphological characters only. Significant differences in six characters were found between species. However, the variation in species markedly overlapped. In our opinion, such external morphological characters could not be used as reliable criteria for species identification (in Mordovia). A female R. esculenta with body size equal to 109 mm was collected. This is the maximum record for the species from European part of the former USSR.

SOME PECULIARITIES OF HUNTING OF WOLVERINE (GULO GULO L.) FOR LARGE UNGULATE ANIMALS IN THE FOREST ZONE OF THE KRASNOYARSK REGION

V.V. Kozhechkin, A.S. Shishikin, I.L. Tumanov

In the article there are given little known facts of specific character of nutrition of/ wolverine in the taiga zone of the region within the bounds of 58°—67° North latitude. There is paid attention to the specific peculiarities of the manducatory apparatus structure of this predator, which supplies its maxilla with very strong squeezing power and strict clutch (in comparison with lynx, wolf, bear and fox). There are described in detail some methods of hunting of this predator for large ungulate animals (Alces alces, Rangifer tarandus) in the given region. There is given the analysis of factors influenced on the effectiveness of quarry by the wolverine of its sacrifices.

THE BARK-BEETLES (COLEOPTERA, SCOLYTIDAE) FROM THE SUBFAMILIES HYLESININAE AND SCOLYTINAE (EXCEPT TRIBUS XYLEBORINI, CRYPHALINI, CORTHYLINI) OF THE WESTERN CAUCASUS

M.Yu. Mandelshtam, N.B. Nikitsky, A.Bibin

Authors have analyzed rich own material of bark-beetles (Coleoptera: Scolytidae) collected in Krasnodar Province of Russia and in Republic of Adygeya as well as collections of Scolytids from these territories deposited in the Zoological Museum of Moscow State University and in the Caucasian Biosphere State Nature Reserve. Five new species of bark-beetles are recorded for the first time for Krasnodar Province. Mediterranean Tomicus destruens (Woll.) has never been previously recorded in the Black Sea coastal countries, including Russia, Georgia and the Caucasus in general. Hylurgus micklitzi Wachtl, Crypturgus mediterraneus Eichh., Pityogenes calcaratus (Eichh.) all are indicated here for fauna of Russia for the first time. Pityogenes trepanatus (Noerdl.) «is recorded here for the Caucasus for the first time. Authors have summarized scattered data on the bark-beetle fauna of the Caucasian Nature Biosphere Reserve — the most important protected area in the Western Caucasus. New distribution data and new host-plants for some bark-beetle species in Russia are provided.

COLLECTIONS OF THE ZOOLOGICAL MUSEUM OF THE MOSCOW LOMONOSSOV STATE UNIVERSITY: CATALOGUE OF THE NAME-BEARING TYPES OF LEPIDOPTERAN TAXA DESCRIBED AFTER THE YEAR 1952

A. V. Sviridov

Catalogue of the type specimens of all butterflies and moths (Lepidoptera), which were deposited in the Zoological Museum of MSU since the first catalogization in 1981, is presented. The new catalogue includes only name-bearing types. Eightly types representing 79 nominal taxa are added to the formerly listed 196 name-bearing types representing 91 nominal taxa. Among these are 70 holotypes, 1 neotype and 3 syntypes of two taxa. Authors of these taxa are 35 researchers in entomology, with 27 of them from Russia, and 11 from other countries, including those from the former USSR. Forty seven of the holotypes belong to the Rhopaloceran taxa (37 nominal subspecies and 10 species — all at first description) and 29 to the Heteroceran taxa (4 subspecies and 25 species). Forty three of the holotypes represent taxa described by professional entomologists from various state institutions and 33 — by amateur collectors. In total name-bearing types of 169 nominal taxa are deposited currently in the museum. The collection of name-bearing types is currently stored separately from other collections in a locked cupboard. The author as curator of the collection addresses to all entomologists to share their knowledge of any omissions from the catalogue to make it more complete in future.

BUTTERFLIES FROM THE FAMILIES SATYRIDAE, NYMPHALIDAE, RIODINIDAE AND LYCAENIDAE (LEPIDOPTERA, RHOPALOCERA) OF THE RIVER BASINS OF OKSU -MURGAB AND PSHART (TADJIKISTAN)

A. V. Sotchivko, L. V. Kaabak

This work presents a faunistic list of the East Pamirs butterflies of Families Satyridae, Nym-phalidae, Riodinidae and Lycaenidae, with comments on biology and origin of taxa. Annual studies in 1993—2002 of the Murgab district fauna allowed to supplement data on the location of populations and the borders of taxa areas.

BREEDING BIOLOGY OF BRAMBLING (FRINGILU MONTIFRINGILLA L.) IN TAIGA OF THE MIDDLE YENISEI

V.Yu. Arkhipov

The breeding biology of the Brambling was studied during 1994—1997 in the floodplain forest of the Yenisei River (Mirnoye field station, Central Siberia, 62°20'N, and 89°00'E). In the study area Brambling occurs in a high density of 68 breeding pairs per 1 km2 (range: 53—78 over four years). In total 42 nests were found, and all these nests were located on trees at 2—12, mean 4,5 meters above the ground. Eggs were laid from late May to early July with two peaks occurring in mid-June and early July. Two of 23 colour-ringed pairs managed to raise two successive broods in one season. Average clutch size was 5,96 eggs (SE ± 0,23, range 4—8). Three of 17 colour-ringed males were proved to be bigynous. None of 55 breeding adults and 85 nestlings ringed during the four years of studies was recorded back to the study area in later years. Thus, neither breeding site tenacity nor philopatry was recorded.

SOME PROBLEMS OF MORPHOGENESIS AND SEASON GROWTH OF THE SHOOTS OF DRABA SIBIRICA (PALL.) THELL.

O.V. Grigoryeva

Morphogenesis of Draba sibirica (Pall.) Thell. has been investigated in several geographically isolated populations. We separated two biomorphs that differ in the shoot length and maintaining the main root. In long-term moist conditions there form shoots of a higher length and, with the length increasing, the plant looses a connection with the main root. Under short-term moist conditions D. sibirica forms relatively small shoots, the connection with the main root being lost. We described the seasonable shoot growth and some peculiarities of the inflorescence and replacing shoots formation inside the buds. During a year there forms one short fragment of a shoot, this easily permitting to detect the age both of separate shoots and the plant as a whole. D. sibirica has apical inflorescence initiated and formed in different time, from the summer beginning and up to the spring. The simultaneous spring flowering occurs due to the slowing down the of the perfectly formed inflorescence growth in the cold period of the year. Just at that time there continues a formation and development of generative organs on the other shoots this leading to an accumulation of inflorescence that simultaneously flower with the temperature rise. The replacing shoots are formed in bosoms of covering leaves neighboring to the inflorescence simultaneously with the formation of its own. They initiate in the early phase of the covering leaves morphogenesis and develop together with them or even faster. The flowering in the end of the summer is one of consequences of a non-synchronous inflorescence initiation.

A.K, Skvortsov. A FEW ADDITIONS TO THE FLORA OF THE SMOLENSK PROVINCE

A.A. Notov, N.R. Markelova, L.V. Kolosova. THE SECOND FINDING OF ASPLENIUM VIRIDE IN MIDDLE RUSSIA

E.L. Zheleznaya. FLORISTIC RECORDS FROM TALDOM DISTRICT OF MOSCOW PROVINCE

A.P. Seregin. ERIGERON DROEBACHIENSIS O.F. MUELL. (COMPOSITAE), A NEW SPECIES FOR MIDDLE RUSSIA

A.K. Skvortsov. MATERIALS TO THE FLORA OF THE KALUGA PROVINCE

A.P. Seregin, Ye.Yu. Zamesova. FLORISTIC RECORDS ON SATINO TRAINING POLYGON (BOROVSK DISTRICT, KALUGA PROVINCE)

T.B. Silaeva, I.V. Kirjukhin. MATERIALS TO THE FLORA FOR SURA RIVER BASIN

L.A. Agafonova, G.A. Kupatadze. NEW AND RARE VASCULAR PLANT SPECIES OF URBANIZED LANDSCAPES OF BELGOROD PROVINCE

V.A. Agafonov, A.P. Sukhorukov. ADDITIONS TO THE FLORA OF VORONEZH PROVINCE