BULLETIN OF MOSCOW SOCIETY OF NATURALISTS

Biological series Volume 107, Part 1 2002

CONTENTS

Triseleva T.A., Safonkin A.F. The structureof the pheromone gland of Archips podana (Scopoli) (Lepidoptera:Tortricidae) and attraction of males by females from different hostplants 3

Schigel D.S. Beetle complexes in polypore fungi in EasteEuropean Plain and Crimea 8 Nikitsky N.B., Tatarinova A.F. Fauna andecology of the minute brown scavenger beetles (Coleoptera, Lafridiidae)of the European North-East of Russia, with the remarks of xylophilous (and some other) beetles from the Moscow region 22

Poltarukha O.P. Diagnostic features and a key for identification of naupliarstages of barnacles (Cirripedia Thoracica) permanently inhabitingthe Black Sea 26

Skvortsov A.K. Some logico-semantic preliminariesto a theory of systematics 32

Khanina L.G., Smirnav V.E., Bobrovsky M.V. Analysis of forest vegetation in the reserve

Kaluzhskie Zasekiby methods of numerical ecology 40

Osipov S. V. Plantsynusiae of the taiga-goltsy landscapes of the Bureinskiy highland(the Russian Far East) 49

Scientific communications

Bolshakov L. V. Complex regional analysis of thebumet moths (Lepidoptera: Zygaenidae) fauna of Tula and neighbouringprovinces 57

Sarycheva E.P. Ontogeny descriptionin three species of the family Umbelliferae that are frequently found in black alder forets of Nerusso-Desnyanskoe polesye 63

Mazurenko M. T. Ontogeny of Laurocerasus officinalis L. (Rosaceae)in Adjaria 70

To the history of science

Nakhapetov B.A. Anapology of I.T. Radozhitsky, a general and botanist 80

Critique and bibliography

Majorov S.R. Review on the book: E.T. Valagina-Malutina "Trees and Shrubs in winter" 88

THE STRUCTURE OF THE PHEROMONE GLAND OF ARCHIPS PODANA (SCOPOLI) (LEPIDOPTERA: TORTRICIDAE) AND ATTRACTION OF MALES BY FEMALES FROM DIFFERENT HOST PLANTS

’.€. Triseleva, A.F. Safonkin

Summary

Thepheromone gland (PG) is located on the dorsal side, under the cuticle,between the 7th and 8th segments of the imago abdomen. On a microscopicslide the PG looks like a loop stretched along the body, which formsnumerous sockets. It consists of one layer of large, tightly packed columnar cells of epithelium. The PG cavity has an external outlet. Thecell nuclei are located closer to the basal membrane. The paternalgenotype influences the PG morphometry. Body mass of a fermale doesnot correlate with morphometry of her PG cells. The development of larvae on an apple-tree, lilac or plum-tree has an inflience upon thelarger diameter of the nucleus and its volume. The volume of the PGnuclei in imagos from a plum-tree was the smallest. These females wereable to attract the least number of males, and males of the ‚ phenotype predominated in this case.

BEETLE COMPLEXES IN POLYPORE FUNGI IN EASTEEUROPEAN PLAIN AND CRIMEA

D.S. Schigel

Summary

Most of beetles,living in polypore fungi are polyphagous. Composition of species livingin widely distributed polypore fungi changes latitude direction,with species diversity and diet spectrum increasing southwards. Beetlelarvae are located mostly in context and in the layer between contextand hymenophore of a fungus. The carpophore accretion leads to changes inthe beetle dimensional distribution. Distribution of beetles inside thecarpophore is constant in different parts of a fungus areal, in spiteof changes in species composition. of beetles. Not only taxo-nomy of fungi determines distribution of beetle larvae, but, also, hardness,structure and hyphal type of carpophore ave important properties ofsubstrate.

FAUNA AND ECOLOGY OF THE MINUTE BROWN SCAVENGER BEETLES(COLEOPTERA, LATRIDIIDAE) OF THE EUROPEAN NORTH-EAST OF RUSSIA, WITHTHE REMARKS OF XYLOPHILOUS (AND SOME OTHER) BEETLES FROM THE MOSCOWREGION

N.B. Nikitsky, A.F. Tatarinova

Summary

DIAGNOSTIC FEATURESAND A KEY FOR IDENTIFICATION OF NAUPLIAR STAGES OF BARNACLES (CIRRIPEDIATHORACICA) PERMANENTLY INHABITING THE BLACK SEA

O.P. Poltarukha

Summary

A comparative analysis of morphology of naupliar stages of all speciesof barnacles (Cirripe-dia Thoracica) permanently inhabiting the BlackSea has been carried ont. Special attention is paid to the features,which are important for of identification larvae. A key for identificationof all naupliar stages has been compiled. The key can be useful also foridentification of the barnacles species from the Caspian and Azov seas,as well as from the Finnish Gulf of the Baltic Sea.

ANALYSIS OF FORESTVEGETATION IN THE RESERVE KALUZHSKIE ZASEKI BY METHODS OF NUMERICALECOLOGY

L.G. Khanina, V.E. Smirnov, M.V. Bobrovsky

Summary

A newclassification procedure was proposed for analysis of forest vegetation oflarge areas (tens of thousands hectares), which is typical for forestryenterprises, natural strict reserves ("zapovedniks") and nationalparks in Russia. Input data of the procedure was species abundancedata at sample plots of fixed area (usually 100 m2). Data for herbs,shrubs, understory and over-story trees were used. The classificationprocedure included severai main stages.

The first stage was preliminaryclassification of sample plots. Database technology was used. Referencedatabase on ecological-coenotic groups developed by A.A.Nitzenko (1969)with addi- tions by O.V.Smimova and L.B.Zaugolnova (unpublished data)was used. For the beginning, all sample plots were divided into largegroups according to dominant tree species in ovcrstory. Then sampleplots within large groups were divided into subgroups by dominance ofecological- coenotic' groups in understory.

At the second stage,preliminary classification of plots was specified. Results of clusteranalysis and ordination of herb species abundance data specified acomposition of subgroups for plots included in the large groups.

Atthe third stage, herb species abundance data for all plots were ordinatedusing detrended correspondence analysis (DCA). For interpretationof the major axes of variation in the vegeta- tion, rank ecologicalcharacteristics of plots were calculated using ecological values of plant species developed by different authors (Ellenberg, 1974; Landolt, 1977,Tsyganov, 1983).

Statistical test of differences among all groupsand subgroups was the last stage of the pro- cedure. Multi-responsepermutation procedures (MRPP, Zimmerman et al., 1985) were used for it.

The algorithm was used for classification of vegetation inthe strict nature reserve Kaluzhskie zaseki (Kaluga region. CentralRussia). Analysis of 755 sample plots with 536 vascular species identified 13 vegetation types. They were as follows: inultispeciesbroad-leaved forest; black-alder forest; spruce forest with nemoralherbs; spruce forest with boreal herbs; pine forest with nemoral herbs; pine forest with grassland-and-edge herbs; pine forest after firewith pure pine-forest herbs; aspen forest; willow communities; birchforest with nemoral herbs; birch forest with grassland- meadow-edgeherbs; mezofite meadows; hygrophyte meadows. All communities differed significantly and their location in the ordination diagram waswell interpreted by the ecological characteristics of plots.

The use of enlarged types of vegetation communities with referencesdatabases and formal methods of data classification simplifies andfacilitates essentially a process of classification of vegetationof large areas. Application of the procedure for Kaluzhskie zasekivegetation has showed success of the ecological-coenotic approachto vegetation classification.

COMPLEX REGIONAL ANALYSIS OF THEBURNET MOTHS (LEPIDOPTERA: ZYGAENIDAE) FAUNA OF TULA AND NEIGHBOURINGPROVINCES

L.V. Bolshakov

Summary

Ecological, chorologo-ecologicaland sozobiological classification of burnet moths (Lepidop-tera: Zygaenidae) in Tula, Moscow and Kaluga provinces of European Russiain suggested. The stude summarizes results of long-term research,collecting and literature sourses on distribution, ecology and biologyof species of regional fauna.

ONTOGENY DESCMPTION OF THREE SPECIESOF FAMILY UMBELLIFERAE THAT ARE FREQUENTLY FOUND IN BLACK ALDERFORESTS OF NERUSSO-DESNYANSKOE POLESYE

E.P. Sarycheva

Summary

This paper maintains description of life stages of several dominants ofherb cover of black alder communities. In ontogeny of each species,we have distinguished following life stages: seedling, juvenile,pre-reproductive adult, reproductive adult, senescent adult. The followingparameters for discribtion of each life stage are used: lenght, widthand number of leaves, roots, stems, number of flowers.

ONTOGENY OFLAUROCERASUS OFFICINALIS L. (ROSACEAE) IN ADJARIA

M.T. Masurenko

Ontogeny and life-forms of (Lauroserasus officinalis L. in South Kolkhida(Adjaria)) were studied. In the lowmountain subtropical belt (from sealevel to 500 meters above sea level) there exist two life-forms - (1)trees, up to 10 meters high, and (2) vegetative movable elfin. Shrub.